Mike Rogers, the chairman of the House intelligence committee, said the spying apparatus had thwarted "dozens" of terrorist plots, details of which he was working to declassify.
The Michigan Republican told CNN that the system was a "lockbox" containing "only phone numbers", rather than recordings. He said it was used "sparingly" and "absolutely overseen by the legislature".
Edward Snowden, the rogue NSA contractor who leaked top secret documents to the media, claimed that even as a low-ranking official he could tap the phones of any American.
Yet intelligence chiefs insist the content of conversations can only be obtained in extraordinary circumstances and with court authority, on the basis of specific evidence of a terrorist plot.
Then, a system called Nucleon records calls, while Prism, which was exposed in documents leaked by Snowden, captures emails, social network messages and other online chats.
William Binney, a former NSA technical director who helped create the agency's eavesdropping network, estimated last week that it records the calls of up to a million people on a "target list".