Appearing via a television link from Los Angeles, Morgan heard a robust intervention from Lord Leveson himself saying he was "perfectly happy" to call Mills and ask her if she had given permission to Morgan to listen to the taped message.
The CNN host then told the inquiry that, during the celebrity couple's bitter divorce, it had been suggested that Heather Mills had recorded telephone messages from her husband and played them to journalists.
Jay repeatedly pressed Morgan about how much he knew, as an editor who had been in charge of two of Britain's largest tabloid titles, about the culture of phone hacking that has often been described as "widespread".
Morgan painted a picture of himself as a detached editor who left the running of his newspaper to other executives, who did not want to know the sources for major stories, and at one stage suggested that editors knew "only 5 per cent of what was going on".
He told the inquiry that, during his years of editing the Daily Mirror Morgan offers tape rumour
between 1995 and 2004, he did not believe that phone hacking was practised. Asked about rumours in Fleet St that the Mirror had been part of a culture of so-called "dark arts", Morgan denied he had "any reason [to believe] it was going on".
Questioned about the evidence the former NOTW editor Rebekah Brooks had given to a parliamentary select committee in which she admitted to occasional payments to police officers, Morgan said that practice, which is illegal, had never happened when he was in charge of the Trinity Mirror paper.
Morgan said private investigators had been used by the Mirror, but said he was "never directly involved" in their commissioning or what they brought in. The Guardian was the self-appointed "bishops of Fleet Street".
- Independent