Newlyweds Ellie and Jon Udall have died of their injuries after a helicopter crash.
A fifth Briton has died from her injuries 16 days after a helicopter crashed in the Grand Canyon.
Ellie Udall, 29, was on honeymoon with her husband Jonathan, 31, and were touring the Grand Canyon when the Airbus EC130 B4 crashed just before sunset on Saturday February 10.
The critically injured couple were being treated in hospital in America, alongside Jennifer Barham, 39, and pilot Scott Booth, 42 but have now died.
Ellie Udall managed to clamber free but her husband was trapped in the wreckage for eight hours as emergency services struggled to get access to the scene of the crash.
Barham and Booth are still in hospital.
Almost £23,000 ($44,000) has been raised for Jonathan and Ellie Udall, according to a JustGiving crowdfunding webpage, according to Chris Tucker, who set up the page.
He wrote: "It deeply saddens me once more to update this page. Yesterday Ellie lost her battle to her injuries, the only consolidation I can muster is that she is no longer in pain and is, I hope, with Jon.
"The continued support to this page is heartwarming during this tragic time. May we never forget this wonderful couple."
Jonathan died four days ago.
On that day Tucker wrote on the JustGiving page: "It is with a very heavy heart that I must type this.
"Our good friend Jon Udall has succumb[ed] to his injuries.
"He was strong, brave and I will never forget him. I will update this page when possible. Ellie is critical and is continuing to fight."
Jonathan Udall's parents Philip and Marlene are in the US.
Traumatised witnesses watched the chopper plummet 180m into the rocky canyon and burst into flames before survivors were seen staggering out of the inferno. One woman was heard screaming.
Friend Rob Fanstone wrote on Facebook: "Words can't describe Jon enough. What a true gentleman he was and touched many, by knowing him from age five, going through primary & secondary schools, going to each other houses after school in primary school.
"The world lost another star and caring person. Life is so cruel at times. You'll sadly be missed by many people."
A neighbour described Udall as "such a lovely bloke, he was very pleasant".
Jonathan Udall, who is originally from Southampton, was a southeast regional manager for Yorkshire Building Society, where he had worked for nearly 10 years.
Becky Dobson, 27, her boyfriend Stuart Hill, 30, and his brother Jason Hill, 32, all died in the crash.
They had also been celebrating Hill's 30th birthday with a trip to Las Vegas.
A British man and his new bride saw the tragic Grand Canyon helicopter crash just minutes after they got married.
Finance worker Chris Parish, from London, exchanged vows with Swedish fiancee Anna Haglund in an intimate ceremony with just the two of them.
But just minutes after the happiest moment of their lives, they watched as the helicopter "crashed and exploded in front of our eyes".
Haglund took to Instagram and posted a picture of the Grand Canyon, together with a broken heart emoji.
She then shared a photo of her and her husband Chris' wedding rings.
The couple's ceremony was held by Las Vegas reverend Lionel Douglass, 63, who
described seeing the horror crash.
He said: "I noticed a helicopter coming into the Grand Canyon and I saw it making a circle and I said to myself, 'that's not where they usually land'.
"I turned to my pilot and asked about the helicopter, and as soon as we turned around and looked at it, it tilted, went down, broke in half, then hit the ground and there was a big explosion."Stuart and Jason's grief-stricken father, Reverend David Hill, said a "light has truly gone out" but took comfort in the "incredibly close" brothers dying together.
He choked back tears as described his sons, Stuart, a Mercedes car salesman, and Jason, a solicitor, as "wonderful boys" who were "inseparable".
The three, who are originally from Worthing, West Sussex, died from multiple injuries, Mohave County Medical Examiner's Office said.
Hundreds of mourners attended a memorial service in Worthing to pay tribute to the victims last week.
Seven candles were lit: three were purple in memory of those who died and four white symbols of hope for the survivors still in hospital.
The helicopter crashed on tribal land in a section of the Grand Canyon outside of the national park where air tours are not as highly regulated.
The National Transportation Safety Board is investigating the crash, and will interview witnesses, survivors, the helicopter operator and manufacturer and others before issuing a full report in at least a year's time.
Preliminary findings are expected before the end of the month.