More than 10,000 athletes from 206 different nations will compete for glory in this year's Olympic Games in Rio. But when it comes to distance running it's likely that, as in previous years, the finals will be dominated by athletes from East African countries or those of East African heritage.
Rio Olympics 2016: East Africans - best by a distance
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Kenya's David Rudisha, right, competes in a men's 800-meter heat during the 2016 Summer Olympics in Rio. Photo / AP
If genetic research alone cannot explain the dominance of East African distance runners, then what other factors might be behind their success? One factor often suggested is the extensive walking and running these athletes undertake from an early age - a total distance run to and from school is often cited at between 5km and 20km. However, this early introduction to endurance training does not appear to result in a higher maximal aerobic capacity (a key determinant of endurance performance) than that seen in elite European distance runners.
Such great heights
What about altitude? Many of the elite Kenyan and Ethiopian distance runners were born and raised at altitudes of around 2000-2500 metres. This may lead to superior levels of haemoglobin (a protein in red blood cells that carries oxygen throughout the body) and haematocrit (the volume of red blood cells in the blood). In turn, this leads to an increased ability to transport oxygen to the working muscles.
While just living at such altitudes may not alone explain the success, it appears that East African athletes also have the ability to train at high-intensity while at altitude. This is something that athletes without continual altitude exposure would find difficult to replicate, so it seems logical to assume that prolonged altitude exposure and the ability to train at a high-intensity while at altitude may contribute to the success of East African distance runners.
A final reason often suggested for the East African dominance of distance running is the motivation to achieve economic success. In relatively poor countries, success in lucrative distance running events can considerably advance an athlete's position in society.
We still can't say conclusively what is behind the phenomenal success of the East African distance runners. But an optimal body type leading to excellent biomechanical efficiency may well play a part, alongside the runners' prolonged exposure to altitude and psychological motivation to succeed.
• Andy Galbraith is a senior lecturer in Exercise Physiology, University of East London