Others theorised that the fishermen likely failed to sever the fish's nerves - which then caused involuntary muscle reflexes.
Muscle motor neurons are triggered by electrical signals, and they can still be active even after the brain has died.
Fishermen in Japan usually prevent this by inserting a spike in the fish's hindbrain, in a method called ikejime.
Some even suggest that the taste of the fish can be affected by failing to carry out ikejime, because the fish's muscles produce lactic acid as they flounder about.