Police have not confirmed the identity of the remains but say they most likely belong to Hollingworth.
"I concluded that the woman was attacked by wild animals who devoured her," Kifnidis said. "I think she was eaten by a wolf or wolves."
Wolves roam remote parts of Greece and have been reported in the Petrota area but attacks on humans are very rare and no fatal attacks have been reported in decades.
Kifnidis said there was a slight possibility that Hollingworth fell victim to a criminal attack by another human, but added "this is unlikely, as she herself phoned her brother and said she was being attacked by dogs."
Tests on animal hairs found among the remains are expected to identify the carnivores, he added.
Police said Hollingworth had been visiting a seaside archaeological site at Messimvria on Thursday, where she arrived by taxi from the village of Maroneia - home to another ancient site - about 30km away by road. She was last seen alive later in the day on a path that forms a shortcut between the two sites.
The remains were found in the hills inland, toward Petrota.
- AP