The latter has back-up in the form of Willem Alberts, to the probable dismay of Wales considering that Albert almost single-handedly changed the course of the game when he came on in the second half in Cardiff last year.
In the front row, Beast Mtawarira has been preferred to last year's SA Rugby Player of the Year, Gurthro Steenkamp, who is part of a powerhouse and very experienced bench with Alberts, Bismarck du Plessis, CJ van der Linde and Johann Muller, who is making his first appearance for the Boks since 2009.
De Villiers has made five changes to the team that defeated New Zealand 18-5 in Port Elizabeth in the Boks' final Tri-Nations match, recalling captain John Smit at hooker, loosehead prop Mtawarira, Burger, and fullback Frans Steyn while Rossouw replaces Botha, who remains in doubt for next week's match against Fiji because of his Achilles tendon problem.
This Springbok team is the most experienced in history with 815 caps - narrowly surpassing the 810 of the team that played Australia in Durban in August.
The only two backline reserves are Butch James and Francois Hougaard, with the latter covering halfback and wing and the former first five-eighths and centre.
With wind and rain forecast, the Boks might have a slight advantage since they are accustomed to the conditions at the Westpac Stadium.
- INDEPENDENT