He said Mr Key today "lost his rag in front of the people of this country and he's spat the dummy in front of the people who've loved him, mainly the media''.
"I think that a leader who has got grit and courage should never do that.
"Above all if he's going to be a bloke that has a beer with the boys and drinks out of the bottle how about being a real bloke and come clean.''
Dr Brash said he didn't care about the content of the recording and had spoken to Mr Banks about them several times.
"He's assured me that we're going into this election together and that I'm the leader.''
He said he would "absolutely'' serve out a three-year term if he made into Parliament and later told reporters he hoped to contest the 2014 election as well.
Mr Harawira also turned in a strong performance, finding considerable ground with his former Maori Party colleague Tariana Turia as the pair championed the rights and welfare of Maori and the poor.
Facing questions on the economy and the prospect of another global financial crisis Mr Harawira got the first round of applause from the studio audience when he said he would be out in the streets if any Government responded to a global recession by cutting benefits. The poor were "bleeding'' already he said.